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38. The few times that they do occur, it is the possessor who tries to make someone leave the circle. Reciprocity can be explored in captivity by handing one chimpanzee a large amount of food, such as a watermelon or leafy branch, and then observing what follows. ( ① ) The owner will be center stage, with a group of others around him or her, soon to be followed by newly formed groups around those who obtained a sizable share, until all food has been distributed. ( ② ) Beggars may complain and cry, but aggressive conflicts are rare. ( ③ ) She will hit them over their head with her branch or bark at them in a high-pitched voice until they leave her alone. ( ④ ) Whatever their rank, possessors control the □□□□□. □□□□□. □□□□□.
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